A regional boss and four other suspected members of the Los Zetas drug cartel were arrested by Mexican marines in the northern state of Coahuila, the Navy Secretariat said Tuesday.

Miguel Angel Rodriguez Diaz was arrested on Sunday along with another man, identified as Olegario Moreno Ibarra, following a shootout with marines in the town of Guerrero, the secretariat said in a statement.

Rodriguez Diaz ran the Zetas cartel's business in northern Coahuila, where he had "operational control over members of the organization in the cities of Piedras Negras, Ciudad Acuña and Cinco Manantiales," the secretariat said.

Marines seized two rifles, two handguns, two rockets, a fragmentation grenade, 1,376 rounds of ammunition, 16 ammunition clips and four packages of cocaine, among other items, from the suspects, the secretariat said.

Two other suspects, identified as Martin Rolando Villa and Oscar Rene Urias Carrillo, were captured by marines in a separate operation in the town of San Carlos on Sunday, the secretariat said.

Urias Carrillo is suspected of being an "accounting assistant for the criminal organization's financial operator," who reported to Rodriguez Diaz, the secretariat said.

Marines seized two handguns, a rifle, $123,920, 2.3 million pesos (about $179,000), 380 rounds of ammunition, ammunition clips and three packages of marijuana from the two men.

A patrol, meanwhile, arrested Jose Odilon Ramirez Perales, who was carrying a suitcase containing a handgun, eight packages of cocaine, more than 460 rounds of ammunition and other items, in the city of Monclova, the secretariat said.

Ramirez Perales told investigators that he belonged to Los Zetas and was in charge of weighing, packaging and shipping different types of drugs.

Heriberto Lazcano Lazcano, the top Los Zetas boss, was killed in a shootout with marines in Coahuila state on Oct. 7.

Lazcano, known as "El Lazca," deserted from the Mexican army in 1999 and formed Los Zetas with three other soldiers, all members of an elite special operations unit, becoming the armed wing of the Gulf drug cartel.

After several years on the payroll of the Gulf cartel, Los Zetas, considered Mexico's most violent criminal organization, went into the drug business on their own account and now control several lucrative territories. EFE